Hoya Australis turning yellow/blotchy (question in comments)

My Hoya Australis has been looking a little mottled for some time now, with a few darker circles on some of the leaves nearer the top.

It’s always looking a bit wrinkled and soft so I upped the watering a little since it’s been warmer.

This was a mistake. The bottom leaves started yellowing, one of which rotted off, another two pulled off pretty easily though weren’t rotten.

Pretty sure I overwatered, but I’m thinking there was something else going on first.

What tends to cause the leaves to be wrinkly and ‘leathery’?

It’s under a grow light where I have a number of other happy plants including Hoyas. I generally don’t have issues with any of my Hoyas so this is perplexing me a bit....

Humidity in this room is usually around 40% which is low but not super low.

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